With some verbs, and in some cases, it will not make much difference whether you use "on" or "onto." However, in cases such as your sentence, "onto" is necessary. Think of it this way: you need the aspect of "on" because walrus are now living "on" this patch of shoreline - they do NOT "live the shoreline." They live ON the shoreline. The aspect of "to" is required because of the movement that took place: They were forced to move FROM one place TO this shoreline.