Both your sentences are the same ;-)
But it should be 'in', not 'during'. The colonies are settlements of people, so are grammatically like cities, towns, villiages, settlements, groups, etc, which would all take 'in'. On the other hand, 'during' can only be used for periods of time. (for periods of time, there ARE cases where either word is acceptable, like "during/in American colonial times" but during can only be used for time periods, where as 'in' has wider uses.